Aims: Protein kinase C epsilon (PKCε) has been shown to play a causative role in the generation of glucose intolerance and insulin resistance. Tissue-specific deletion of PKCε indicates multiple sites of action, further validating the kinase as a potential target for the treatment of T2D. Here, we examined the effects of a recently-developed PKCε inhibitor (CIDD-0150612) on insulin signalling in palmitate-treated HepG2 hepatocytes and also acute and long-term effects on glucose homeostasis in fat-fed mice.
Methods: HepG2 cells were treated for 18 h with 500 µM palmitate and 10 µM CIDD-0150612, and stimulated with 30 nM insulin for 10 min after a 2 h serum-free period. Mice were fed a high-fat diet for 8 weeks and treated with CIDD-0150612 (40 mg/kg ip eod) in the final 2 weeks.
Results: In contrast to reports that PKCε mediates inhibitory threonine phosphorylation of the insulin receptor (IR), CIDD-0150612 had no effect on insulin-stimulated IR tyrosine phosphorylation in lipid-treated hepatocytes, nor on downstream IRS-1 tyrosine phosphorylation. However, the inhibitor promoted Akt phosphorylation in a highly insulin-dependent manner, and reversed the 40% inhibition of insulin-stimulated Akt phosphorylation by palmitate (P<0.01). Fat-fed mice treated with CIDD-0150612 had reduced body fat (14.6 vs 20.2%, P<0.01) and body weight (25.4 vs 28.3 g, P<0.001) compared to vehicle-treated littermates. Mice treated acutely with CIDD-0150612 exhibited elevated fasting blood glucose (15.7 vs 10.0 mM, P<0.05). However, mice subjected to glucose tolerance tests 24h after the last inhibitor dose had lower fasting glucose (8.6 vs 10.3 mM, P<0.05), improved glucose tolerance (AUC 1162 vs 1626 mM.min, P<0.001) and lower insulin excursions (AUC 49 vs 88 ng/ml.min, P<0.05).
Conclusions: The PKCε inhibitor CIDD-0150612 had beneficial effects on insulin signalling, fat mass and glucose homeostasis. Because certain effects were not previously observed in studies of PKCε-deficient mice, off-target effects may be partly responsible.